Undergraduate education

Duke University, BS, 1989

Years in MSTP



Biochemistry, 1999


Judith Kimble, PhD

Thesis title

Control of Organ Shape by the GON-1 Metalloprotease


Pathology, Brigham and Women’s Hospital, 2001-2003


Transfusion Medicine, Harvard, 2002-2004

Current position

Professor, Department of Urology, University of California-San Francisco, School of Medicine

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Undergraduate and/or masters publications

Reznikoff CA, Belair CD, Yeager TR, Savelieva E, Blelloch RH, Puthenveettil JA, Cuthill S. A molecular genetic model of human bladder cancer pathogenesis. Semin Oncol. 1996 Oct;23(5):571-84. PMID: 8893868.

PhD publications

Mathies LD, Schvarzstein M, Morphy KM, Blelloch R, Spence AM, Kimble J. TRA-1/GLI controls development of somatic gonadal precursors in C. elegans. Development. 2004 Sep;131(17):4333-43. PMID: 15294864.

Blelloch R, Anna-Arriola SS, Gao D, Li Y, Hodgkin J, Kimble J. The gon-1 gene is required for gonadal morphogenesis in Caenorhabditis elegans. Dev Biol. 1999 Dec 1;216(1):382-93. PMID: 10588887.

Blelloch R, Newman C, Kimble J. Control of cell migration during Caenorhabditis elegans development. Curr Opin Cell Biol. 1999 Oct;11(5):608-13. PMID: 10508660.

Blelloch R, Kimble J. Control of organ shape by a secreted metalloprotease in the nematode Caenorhabditis elegans. Nature. 1999 Jun 10;399(6736):586-90. PMID: 10376599.

Roehl H, Bosenberg M, Blelloch R, Kimble J. Roles of the RAM and ANK domains in signaling by the C. elegans GLP-1 receptor. EMBO J. 1996 Dec 16;15(24):7002-12. PMID: 9003776; PMCID: PMC452526.