Undergraduate education

Vanderbilt University, BA, 2009

Years in MSTP



Cellular and molecular biology, 2015


Dan Greenspan, PhD

Thesis title

Multifaceted Roles of Type V Collagen: Fibrillogenesis in Embryonic Development, Ehlers-Danlos Syndrome, and Autoimmunity in Atherosclerosis


Internal medicine, Washington University, School of Medicine, 2017-

Current position

Resident, Department of Medicine, Washington University, School of Medicine

PhD publications

Sullivan JA, Jankowska-Gan E, Hegde S, Pestrak MA, Agashe VV, Park AC, Brown ME, Kernien JF, Wilkes DS, Kaufman DB, Greenspan DS, Burlingham WJ. Th17 responses to Col V, kα-1-tubulin and vimentin are present early in human development and persist lifelong. Am J Transplant. 2017 Apr;17(4):944-956. Epub 2016 Dec 19. PMID: 27801552; PMCID: PMC5626015.

Park AC, Huang G, Jankowska-Gan E, Massoudi D, Kernien JF, Vignali DA, Sullivan JA, Wilkes DS, Burlingham WJ, Greenspan DS. Mucosal administration of collagen V ameliorates atherosclerotic plaque burden by inducing IL-35-dependent tolerance. J Biol Chem. 2016 Feb 12:291(7):3359-70. PMID: 26721885; PMCID: PMC4751380.

Yang C, Park AC, Davis NA, Russell JD, Kim B, Brand DD, Lawrence MJ, Ge Y, Westphall MS, Coon JJ, Greenspan DS. Comprehensive Mass Spectrometric Mapping of the Hydroxylated Amino Acid residues of the α1(V) Collagen Chain. J Biol Chem. 2012 Nov 23;287(48):40598-610. PMID: 23060441; PMCID: PMC3504773.

Park AC, Phillips CL, Pfeiffer FM, Roenneburg DA, Kernien JF, Adams SM, Davidson JM, Birk DE, Greenspan DS. Homozygosity and Heterozygosity for Null Col5a2 Alleles Produce Embryonic Lethality and a Novel Classic Ehlers-Danlos Syndrome-Related Phenotype. Am J Pathol. 2015 Jul;185(7):2000-2011. PMID: 25987251; PMCID: PMC4483463.

Park AC, Phan N, Massoudi D, Liu Z, Kernien JF, Adams SM, Davidson JM, Birk DE, Liu B, Greenspan DS. Deficits in Col5a2 Expression Result in Novel Skin and Adipose Abnormalities and Predisposition to Aortic Aneurysms and Dissections. Am J Pathol. 2017 Oct;187(10):2300-2311. PMID: 28734943; PMCID: PMC5809516.