Undergraduate education

Indiana University-Bloomington, BS, 2000

Years in MSTP



Biochemistry, 2007


Alan Attie, PhD

Thesis title

A Novel Role for Cholecystokinin in Beta-Cell Growth and Survival


Pathology, University of Pennsylvania, 2009-2013


Surgical pathology, Oregon Health & Science University, 2013-2014

Hematopathology, Oregon Health & Science University, 2014-2015

Current position

Assistant professor, Department of Pathology, Oregon Health & Science University, School of Medicine

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Undergraduate or masters publications

Raess PW, Mubarak MS, Ischay MA, Foley MP, Jennermann TB, Raghavachari K, Peters DG. Catalytic reduction of 1-iodooctane by nickel(I) salen electrogenerated at carbon cathodes in dimethylformamide: Effects of added proton donors and a mechanism involving both metal- and ligand-centered one-electron reduction of nickel(II) salen. Journal of Electroanalytical Chemistry. 2007 May;603(1):124-134.

PhD publications

Lavine JA, Raess PW, Stapleton DS, Rabaglia ME, Suhonen JI, Schueler KL, Koltes JE, Dawson JA, Yandell BS, Samuelson LC, Beinfeld MC, Davis DB, Hellerstein MK, Keller MP, Attie AD. Cholecystokinin is up-regulated in obese mouse islets and expands beta-cell mass by increasing beta-cell survival. Endocrinology. 2010 Aug;151(8):3577-88. PMID: 20534724; PMCID: PMC2940525.

Lavine JA, Raess PW, Davis DB, Rabaglia ME, Presley BK, Keller MP, Beinfeld MC, Kopin AS, Newgard CB, Attie AD. Contamination with E1A-positive wild-type adenovirus accounts for species-specific stimulation of islet cell proliferation by CCK: a cautionary note. Mol Endocrinol. 2010 Feb;24(2):464-7. PMID: 20081104; PMCID: PMC2817600.

Clee SM, Yandell BS, Schueler KM, Rabaglia ME, Richards OC, Raines SM, Kabara EA, Klass DM, Mui ET, Stapleton DS, Gray-Keller MP, Young MB, Stoehr JP, Lan H, Boronenkov I, Raess PW, Flowers MT, Attie AD. Positional cloning of Sorcs1, a type 2 diabetes quantitative trait locus. Nat Genet. 2006 Jun;38(6):688-93. PMID: 16682971.