Undergraduate education

Massachusetts Institute of Technology, BS, 1995

Years in MSTP



Cellular and molecular biology, 2002


Alan Attie, PhD

Thesis title

A Genetic Dissection of Obesity and Type 2 Diabetes Mellitus in the Mouse


Medicine, Yale University, 2004-2009


Endocrinology, Oregon Health and Science University, 2009-2012

Current position

Clinician, Department of Endocrinology, Virginia Mason Medical Center

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PhD publications

Bhatnagar S, Oler AT, Rabaglia ME, Stapleton DS, Schueler KL, Truchan NA, Worzella SL, Stoehr JP, Clee SM, Yandell BS, Keller MP, Thurmond DC, Attie AD. Positional cloning of a type 2 diabetes quantitative trait locus; tomosyn-2, a negative regulator of insulin secretion. PLoS Genet. 2011 Oct;7(10):e1002323. PMID: 21998599; PMCID: PMC3188574.

Clee SM, Yandell BS, Schueler KM, Rabaglia ME, Richards OC, Raines SM, Kabara EA, Klass DM, Mui ET, Stapleton DS, Gray-Keller MP, Young MB, Stoehr JP, Lan H, Boronenkov I, Raess PW, Flowers MT, Attie AD. Positional cloning of Sorcs1, a type 2 diabetes quantitative trait locus. Nat Genet. 2006 Jun;38(6):688-93. PMID: 16682971.

Stoehr JP, Byers JE, Clee SM, Lan H, Boronenkov IV, Schueler KL, Yandell BS, Attie AD. Identification of major quantitative trait loci controlling body weight variation in ob/ob mice. Diabetes. 2004 Jan;53(1):245-9. PMID: 14693723.

Lan H, Stoehr JP, Nadler ST, Schueler KL, Yandell BS, Attie AD. Dimension reduction for mapping mRNA abundance as quantitative traits. Genetics. 2003 Aug;164(4):1607-14. PMID: 12930764; PMCID: PMC1462655.

Lan H, Rabaglia ME, Stoehr JP, Nadler ST, Schueler KL, Zou F, Yandell BS, Attie AD. Gene expression profiles of nondiabetic and diabetic obese mice suggest a role of hepatic lipogenic capacity in diabetes susceptibility. Diabetes. 2003 Mar;52(3):688-700. PMID: 12606510. 

Attie AD, Krauss RM, Gray-Keller MP, Brownlie A, Miyazaki M, Kastelein JJ, Lusis AJ, Stalenhoef AF, Stoehr JP, Hayden MR, Ntambi JM. Relationship between stearoyl-CoA desaturase activity and plasma triglycerides in human and mouse hypertriglyceridemia. J Lipid Res. 2002 Nov;43(11):1899-907. PMID: 12401889.

Ntambi JM, Miyazaki M, Stoehr JP, Lan H, Kendziorski CM, Yandell BS, Song Y, Cohen P, Friedman JM, Attie AD. Loss of stearoyl-CoA desaturase-1 function protects mice against adiposity. Proc Natl Acad Sci U S A. 2002 Aug 20;99(17):11482-6. PMID: 12177411; PMCID: PMC123282.

Nadler ST, Stoehr JP, Rabaglia ME, Schueler KL, Birnbaum MJ, Attie AD. Normal Akt/PKB with reduced PI3K activation in insulin-resistant mice. Am J Physiol Endocrinol Metab. 2001 Dec;281(6):E1249-54. PMID: 11701440.

Stoehr JP, Nadler ST, Schueler KL, Rabaglia ME, Yandell BS, Metz SA, Attie AD. Genetic obesity unmasks nonlinear interactions between murine type 2 diabetes susceptibility loci. Diabetes. 2000 Nov;49(11):1946-54. PMID: 11078464.

Nadler ST, Stoehr JP, Schueler KL, Tanimoto G, Yandell BS, Attie AD. The expression of adipogenic genes is decreased in obesity and diabetes mellitus. Proc Natl Acad Sci U S A. 2000 Oct 10;97(21):11371-6. PMID: 11027337; PMCID: PMC17207.